Following is a table synthesizing published cumulative molecular knowledge on Killifishes.
It concerns only those publications with a systematic purpose and focus at the species group level at least (publications to separate species or to separate groups of populations, e.g. for Aphanius fasciatus, Fundulus parvipinnis, Fundulus zebrinus-kansae, Fundulus heteroclitus-macrolepidotus, Fundulus bermudae-relictus, Aphanius iberus-baeticus, the Chichancanab species flock in Cyprinodon, are not considered) 


By nature, Killi-Data online is constantly revised and this table will be updated and freely available on the website. 



Let's not forget, though, to congratulate those researchers, listed hereafter (full bibliographic references are given after the table), who have brought us to the present state of the art : their work could be erroneously seen as easy with automatic machines… this is not the case since molecular studies require expertise to collect tissues and perform replications of primers, savoir-faire to align sequences and analytical skills to scrutiny the trees obtained (and choose the right software and method of producing them). 



The following list has been updated on October 12. 2016, with nearly 1 200 species/populations

You may save this file as a private copy, and because it is important to researchers, you may also publish this precise file, either printed or on electronic material provided that you do not change anything in it, that you mention your source "Huber, J.H., "Cumulative molecular knowledge on Killifishes" and date of update, © Huber, Killi-Data online <>" and that in any case you use the last update of this file.
Please take into account copyright conditions (see OPTIMIZATION), if you use it.

legend Beware, the automatic translator reverses scientific names : please use the English version to save this list.

KDI-AKA0001AustrolebiasAustrolebias nigripinnis RK 99Howard Wu, USAfish, 100% ethanol
KDI-AKA0002NothobranchiusNothobranchius wattersi Hoba MZMW 09-6Howard Wu, USAfish, 100% ethanol
KDI-AKA0003RivulusRivulus xiphidius Crique Blanche SD-FG 03-10Jack Heller, USAfish, 100% ethanol
KDI-AKA0004NothobranchiusNothobranchius jubbi ASTom Kean, USAfish, 100% ethanol
KDI-AKA0005NothobranchiusNothobranchius wattersi Hoba MZMW 09-6Tom Kean, USAfish, 100% ethanol
KDI-AKA0006SimpsonichthysSimpsonichthys heloplites AS (Syn_ antenori)Howard Wu, USAfish, 100% ethanol
KDI-AKA0007PterolebiasPterolebias luelingi BOL HV-2015-07Howard Wu, USAfish, 100% ethanol
KDI-AKA0008GnatholebiasGnatholebias zonatus Las Mercedes VEN 2014-1Howard Wu, USAfish, 100% ethanol
KDI-AKA0009NothobranchiusNothobranchius hassoni DRCH 2008-10Tom Kean, USAfish, 100% ethanol
KDI-AKA0010NothobranchiusNothobranchius polli DRCH 2008-2Tom Kean, USAfish, 100% ethanol
KDI-AKA0011AphyosemionAphyosemion wachtersi FCO 2011-4Tom Kean, USAfish, 100% ethanol
KDI-AKA0012SimpsonichthysSimpsonichthys heloplites AS (Syn_ antenori)Tom Kean, USAfish, 100% ethanol
KDI-AKA0013NothobranchiusNothobranchius fuscotaeniatus Kitonga TAN 97-9Tom Kean, USAfish, 100% ethanol
KDI-AKA0014AphyosemionAphyosemion punctatum Bolo BaiHoward Wu, USAfish, 100% ethanol
KDI-AKA0015ArchiaphyosemionArchiaphyosemion guineense Lenghe-Curoh SL 93-37Thomas Milkuhn, USAfish, 100% ethanol
KDI-AKA0016FundulopanchaxFundulopanchax deltaensis ASTyrone Genade, USAfish, 100% ethanol
KDI-AKA0017CyprinodonCyprinodon nazas La ConchaJohn Brill, USAfish, 100% ethanol
KDI-AKA0018CyprinodonCyprinodon albivelis 5 km South of GaleanaJohn Brill, USAfish, 100% ethanol
KDI-AKA0019NothobranchiusNothobranchius guentheri ZAN 2014-2Tyrone Genade, USAfish, 100% ethanol
KDI-AKA0020KryptolebiasKryptolebias sp. Guadeloupe Bon Goût GT-15Tyrone Genade, USAfish, 100% ethanol
KDI-FV0001AphyosemionAphyosemion punctatum Buang Bai Lester McCathie, UKfish, 100% ethanol
KDI-FV0002Archiaphyosemion (Nimbapanchax)Archiaphyosemion viride Village Bassalow GM 09/20Lester McCathie, UKfish, 100% ethanol
KDI-FV0003AustrolebiasAustrolebias alexandri El Pingo Lester McCathie, UKfish, 100% ethanol
KDI-FV0004CallopanchaxCallopanchax monroviae (red) Paynesville, Liberia L97 Lester McCathie, UKfish, 100% ethanol
KDI-FV0005CallopanchaxCallopanchax occidentalis (blue) Malai SL03/11 Lester McCathie, UKfish, 100% ethanol
KDI-FV0006Aphyosemion (Chromaphyosemion)Aphyosemion bivittatum (red) Lagos Lester McCathie, UKfish, 100% ethanol
KDI-FV0007FundulopanchaxFundulopanchax gresensi Takwai Lester McCathie, UKfish, 100% ethanol
KDI-FV0008FundulopanchaxFundulopanchax spoorenbergi Lester McCathie, UKfish, 100% ethanol
KDI-FV0009PapiliolebiasPapiliolebias francescae La Fiada BPB 2015/18Lester McCathie, UKfish, 100% ethanol
KDI-FV0010SimpsonichthysSimpsonichthys filamentosus ABPV 13/3Lester McCathie, UKfish, 100% ethanol
KDI-FV0011Simpsonichthys (Hypsolebias)Simpsonichthys shibattai Bom Jesus da Lapa Maikel Meijer, The Netherlandsfish, 100% ethanol
KDI-FV0012RivulusRivulus agilae St. Laurent du Maroni FG 15-01Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0013RivulusRivulus agilae Malali GUY 15-05Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0014RivulusRivulus beniensis Itenez Dpt., Road Bella Vista to Magdalena, Bolivia HV 2015-03Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0015RivulusRivulus berovidesi Salta de Soroa, Cuba Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0016RivulusRivulus chucunaque PAN 2012-01Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0017RivulusRivulus frenatus Anarico, Demerare Distr., Guyana (Br) Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0018RivulusRivulus geayi (Granite stones) Savanne Roche, French Guyane F.Guiane 15-14Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0019RivulusRivulus ornatus Neue Horizonte, Peru Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0020RivulusRivulus speciosus Quista Cocha, Peru Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0021RivulusRivulus tenuis Papalapoan, Mexico Wim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0022RivulusRivulus xiphidius FB 95-02, F. GuyaneWim Suijker, The Netherlandsfish, 100% ethanol
KDI-FV0023AphyosemionAphyosemion caudofasciatum FCO 2011-8Bob Merritt, UKfish, 100% ethanol
KDI-FV0024AphyosemionAphyosemion celiae celiae Makanje Plantation ABK 2007-146Bob Merritt, UKfish, 100% ethanol
KDI-FV0025AphyosemionAphyosemion hanneloreae GEB 94-20Bob Merritt, UKfish, 100% ethanol
KDI-FV0026AphyosemionAphyosemion ogoense pyrophore Komono yellow Bob Merritt, UKfish, 100% ethanol
KDI-FV0027AphyosemionAphyosemion primigenium GBN 88-10Bob Merritt, UKfish, 100% ethanol
KDI-FV0028AphyosemionAphyosemion wachtersi FCO 2011-4Bob Merritt, UKfish, 100% ethanol
KDI-FV0029Aphyosemion (Chromaphyosemion)Aphyosemion volcanum Mile 4 JVC 2005Bob Merritt, UKfish, 100% ethanol
KDI-FV0030FundulopanchaxFundulopanchax amieti Sud Sanaga Bob Merritt, UKfish, 100% ethanol
KDI-FV0031FundulopanchaxFundulopanchax amieti km 18 Edea-Yaounde C89-31Bob Merritt, UKfish, 100% ethanol
KDI-FV0032AustrolebiasAustrolebias nigripinnis De Carmelo Paul Carter, UKfish, 100% ethanol
KDI-FV0033FundulopanchaxFundulopanchax avichang GEML 2000-16Paul Carter, UKfish, 100% ethanol
KDI-FV0034FundulopanchaxFundulopanchax nigerianus Jos Plateau Paul Carter, UKfish, 100% ethanol
KDI-FV0035NothobranchiusNothobranchius cardinalis Lissingi River TAN 1997-27Paul Carter, UKfish, 100% ethanol
KDI-FV0036NothobranchiusNothobranchius flammicomantis FTZ 10-21Paul Carter, UKfish, 100% ethanol
KDI-FV0037ScriptaphyosemionScriptaphyosemion guignardi Mamou Paul Carter, UKfish, 100% ethanol
KDI-FV0038NothobranchiusNothobranchius flagrans Kabungu River CD 13/7Trevor Wood, UKfish, 100% ethanol
KDI-FV0039NothobranchiusNothobranchius guentheri Zanzibar Trevor Wood, UKfish, 100% ethanol
KDI-FV0040NothobranchiusNothobranchius kilomberoensis Ifakara TAN 95/4Trevor Wood, UKfish, 100% ethanol
KDI-FV0041NothobranchiusNothobranchius krysanovi Quelimane MZCS 209-249Trevor Wood, UKfish, 100% ethanol
KDI-FV0042NothobranchiusNothobranchius patrizii Msumalini Creek K96-23Trevor Wood, UKfish, 100% ethanol
KDI-FV0043NothobranchiusNothobranchius rachovii Beira Beira 98Trevor Wood, UKfish, 100% ethanol
KDI-FV0044AphaniusAphanius mento Bor TUBCD 05/17J. Cristiano, Portugalfish, 100% ethanol
KDI-FV0045LeptopanchaxLeptopanchax minimus Campo Grande CERS 01-12Steven MacDonald, UKfish, 100% ethanol
KDI-FV0046AphyosemionAphyosemion ogoense GHP 80-24, GabonRoger Shiner, UKfish, 100% ethanol
KDI-FV0047GnatholebiasGnatholebias zonatus Villavicencio, ColombiaRoger Shiner, UKfish, 100% ethanol
KDI-FV0048NothobranchiusNothobranchius eggersi Rufiji River Roger Shiner, UKfish, 100% ethanol
KDI-FV0049Simpsonichthys (Hypsolebias)Simpsonichthys fasciatus Roger Shiner, UKfish, 100% ethanol
KDI-FV0050Simpsonichthys (Spectrolebias)Simpsonichthys reticulatus Altamira, Rio Xingu, Brasil Roger Shiner, UKfish, 100% ethanol
KDI-FV0051AphyosemionAphyosemion exiguum Moyenam ADK 10-305Eduardo Garcia Lastra, Espanafish, 100% ethanol
KDI-FV0052NothobranchiusNothobranchius rachovii Beira 98 Frans Vermeulen, Arubafish, 100% ethanol
KDI-FV0053AphyosemionAphyosemion calliurum CBL 01-20Roger Gladwell, UKfish, 100% ethanol
KDI-FV0054AphyosemionAphyosemion ogoense pyrophore RPC 78-18Roger Gladwell, UKfish, 100% ethanol
KDI-FV0055AphyosemionAphyosemion striatum GEMHS 00-03Roger Gladwell, UKfish, 100% ethanol
KDI-FV0056Simpsonichthys (Hypsolebias)Simpsonichthys coamazonicus Parnaiba, BrasilFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0057NotholebiasNotholebias minimus CERS 01-12 Campo Grande, BrasilFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0058PapiliolebiasPapiliolebias ashleyae BOL 2015-14, BoliviaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0059RachoviaRachovia hummelincki Cerro Atravesado, Paraguana, VenezuelaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0060RachoviaRachovia sp. "Monteria", near village Monteria, Sinu area, West ColombiaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0061RenovaRenova oscari VEN 2014-07 Isla Raton, Finka Rosalia, VenezuelaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0062RivulusRivulus aff. beniensis BOL 2015-02, roadside ditches Magdalena - BellavistaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0063RivulusRivulus aff. beniensis Lago SuarezFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0064RivulusRivulus aff. limoncochae COL 2014-02 Puerto Gaitan, Meta Distr., ColombiaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0065RivulusRivulus aff. rectocaudatus ICE 2015-22 Calderon trail, Leticia, ColombiaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0066RivulusRivulus aff. rubrolineatus ICE 2015-21 roadside ditches at KM 13 Leticia - CalderonFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0067RivulusRivulus aff. urophthalmus Sipaliwini River, Kwamalasamutu, South West SurinameFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0068RivulusRivulus fransvermeuleni upper Nickery River, road to the West. SurinameFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0069RivulusRivulus leticia ICE 2015-20 KM 13 rd. Leticia to Calderon. In backyard home Fred Vosse.Frans Vermeulen, Arubafish, 100% ethanol
KDI-FV0070RivulusRivulus aff. ornatus Neue Horizonte, PeruFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0071RivulusRivulus pivijay COL 2009-28 Road Media Luna - Pivijay, ColombiaFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0072RivulusRivulus species "Curillo", ICE 2015 Village at upper Caqueta RiverFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0073RivulusRivulus species "Mocoa", ICE 2015 Near Mocoa - Mocoa river, upper Caqueta RiverFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0074RivulusRivulus speciosus ASFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0075Simpsonichthys (Simpsonichthys)Simpsonichthys margaritatus Rio Verde, BrasilFrans Vermeulen, Arubafish, 100% ethanol
KDI-FV0076Simpsonichthys (Spectrolebias)Simpsonichthys brousseaui BOL 2015-14, BoliviaFrans Vermeulen, Arubafish, 100% ethanol
KDI-AKA0021NothobranchiusNothobranchius furzeri GonarezhouTyrone Genadefish, 100% ethanol
KDI-AKA0022NothobranchiusNothobranchius furzeri Massingir MZCS 12-2Tyrone Genadefish, 100% ethanol
KDI-AKA0023NothobranchiusNothobranchius krysanovi MZCS 209-249 QuelimaneTyrone Genadefish, 100% ethanol
KDI-AKA0024NothobranchiusNothobranchius pienaari Lower Limpopo MZCS 2009-201Tyrone Genadefish, 100% ethanol
KDI-KCF0001Simpsonichthys (Spectrolebias)Simpsonichthys pilleti Quimome BFB 12-9  (male and female)Didier Pillet, Francefish, 100% ethanol
KDI-KCF0002Simpsonichthys (Spectrolebias)Simpsonichthys semiocellatus (single fish)Didier Pillet, Francefish, 100% ethanol
KDI-KCF0003AustrolebiasAustrolebias accorsii Terra TypicaDidier Pillet, Francefish, 100% ethanol
KDI-KCF0004Simpsonichthys (Spectrolebias)Simpsonichthys brousseaui El Tinto BFB 12-8Didier Pillet, Francefish, 100% ethanol
KDI-KCF0005AustrolebiasAustrolebias elongatus ARG Ruta11 MP 11-01 (single fish)Didier Pillet, Francefish, 100% ethanol
KDI-KCF0006MaratecoaraMaratecoara lacortei Agua Boa HVP 7-16 (male & female)Didier Pillet, Francefish, 100% ethanol
KDI-SEK0001PronothobranchiusPronothobranchius seymouri GHALOZ 03-16 (6 km N from Kasseh)Amer Faour, Espanafish, 100% ethanol
KD-PL0001AphyosemionAphyosemion ocellatum Sika Sika GJS 00-05Antoni Chwialkowski, Polandfish, 100% ethanol
KD-PL0002ScriptaphyosemionScriptaphyosemion cauveti KindiaAntoni Chwialkowski, Polandfish, 100% ethanol
KD-PL0003AphyosemionAphyosemion bualanum Batibo ADL 13-22Antoni Chwialkowski, Polandfish, 100% ethanol
KD-PL0004Aphyosemion (Diapteron)Aphyosemion georgiae GWW 86-2Antoni Chwialkowski, Polandfish, 100% ethanol
KDI-AKA0025AphyosemionAphyosemion joergenscheeli GJS 00-13Jeff Wasley, USAfish, DSSE
KDI-AKA0026AphyosemionAphyosemion raddai ModeJeff Wasley, USAfish, DSSE
KDI-AKA0027AphyosemionAphyosemion bualanum NtuiJeff Wasley, USAfish, DSSE
KDI-AKA0028AphyosemionAphyosemion polli CIJeff Wasley, USAfish, DSSE
KDI-AKA0029Simpsonichthys (Spectrolebias)Simpsonichthys brousseaui BP 13-7 QuimoneJeff Wasley, USAfish, DSSE
KDI-AKA0030AphyosemionAphyosemion hera GJS 00-29 BenguieJeff Wasley, USAfish, DSSE
KDI-AKA0031ArchiaphyosemionArchiaphyosemion petersi Banco ParkJeff Wasley, USAfish, DSSE
KDI-AKA0032RivulusRivulus waimacui GUY 02-4Jeff Wasley, USAfish, DSSE
KDI-AKA0033AphyosemionAphyosemion sp. aff. christyi Lobaye Lou Hersche, USAfish, DSSE
KDI-AKA0034FundulopanchaxFundulopanchax sjoestedti US aquarium strainLou Hersche, USAfish, DSSE
KDI-AKA0035AphyosemionAphyosemion louessense RPC 78/33Lou Hersche, USAfish, DSSE
KDI-AKA0036AphyosemionAphyosemion bualanum N'TuiLou Hersche, USAfish, DSSE
KDI-AKA0037AphyosemionAphyosemion georgiae GWW 86-2Lou Hersche, USAfish, DSSE
KDI-AKA0038AphyosemionAphyosemion sp. aff. castaneum OyoLou Hersche, USAfish, DSSE
KDI-AKA0039AphyosemionAphyosemion sp. COFE 2010-20Lou Hersche, USAfish, DSSE
KDI-AKA0040AphyosemionAphyosemion labarrei MadimbaLou Hersche, USAfish, DSSE
KDI-AKA0041AphyosemionAphyosemion musafiri AVD 2007-1Lou Hersche, USAfish, DSSE
KDI-AKA0042NothobranchiusNothobranchius korthausae Yellow Aquarium StrainJim Kostich (WAKO, AKA), USAfish DSSE
KDI-AKA0043LamprichthysLamprichthys tanganicanusJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0044NothobranchiusNothobranchius orthonotus Mabalane MZG 06-3Jim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0045AphaniusAphanius mento Ereli Konya, TurkeyJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0046FundulopanchaxFundulopanchax sjoestedti LoeJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0047AphaniusAphanius farsicus Lake MaharluJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0048Simpsonichthys (Simpsonichthys)Simpsonichtys santanae Aquarium StrainJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0049FundulopanchaxFundulopanchax sjoestedti Funge ADL 13-39Jim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0050NothobranchiusNothobranchius kilomberoensis TAN 95/4 IfakaraJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0051FundulopanchaxFundulopanchax filamentosus Majidun NG MM 14Jim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0052FundulopanchaxFundulopanchax sjoestedti Dwarf RedJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0053Simpsonichthys (Opthalmolebias)Simpsonichthys (Opthalmolebias) constanciae Barra de Sao JoaoJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0054FundulopanchaxFundulopanchax sjoestedti Funge ADK 11-432Jim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0055AustrolebiasAustrolebias nigripinnis Villa SorianoJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0056FundulopanchaxFundulopanchax sjoestedti Niger DeltaJim Kostich (WAKO, AKA)fish DSSE
KDI-AKA0057PterolebiasPterolebias longipinnis [aff.] BOL HV 2015-05 Laguna SuarezJim Kostich (WAKO, AKA)fish DSSE


Bibliographic references [in chronological order].

Bernardi, G., P. Sordino & D.A. Powers. 1993. Concordant mitochondrial and nuclear DNA Phylogenies for Populations of the teleost fish Fundulus heteroclitus. Proc. Natl. Acad. Sci. USA, 90 (20): 9271-9274. 

Murphy, W.J. & G.E. Collier. 1997. A Molecular Phylogeny for Aplocheiloid Fishes (Atherinomorpha, Cyprinodontiformes): The role of Vicariance and the Origins of Annualism. Mol. Biol. Evol., 14 (8): 790-799, 6 figs., 2 tabs.  

Murphy, W.J., T.N.P. Nguyen, E.B. Taylor & G.E. Collier. 1999. Mitochondrial DNA Phylogeny of West African Aplocheiloid Killifishes (Cyprinodontiformes, Aplocheilidae). Molecular Phylogenetics and Evolution, 11 (3) (April): 343-350, 3 figs. 

Murphy, W.J., J.E. Thomerson & G.E. Collier. 1999. Phylogeny of the Neotropical Killifish Family Rivulidae (Cyprinodontiformes, Aplocheiloidei) Inferred from Mitochondrial DNA Sequences. Molecular Phylogenetics and Evolution, 13 (2) (November): 289-301, 5 figs. 

Hrbek, T. & A. Larson. 1999. The Evolution of Diapause in the Killifish Family Rivulidae (Atherinomorpha, Cyprinodontiformes): a molecular phylogenetic and biogeographic Perspective. Evolution, 53: 1200-1216., 7 figs., 3 tabs. 

Hrbek, T., F. Küçük, T. Frickley, K.N. Stölting, R.H. Wildekamp & A. Meyer. 2002. Molecular Phylogeny and historical Biogeography of the Aphanius (Pisces, Cyprinodontiformes) species Complex of central Anatolia, Turkey. Molecular Phylogenetics and Evolution, 25: 125-137, 5 figs., 2 tabs.  

Hrbek, T. & A. Meyer. 2003. Closing of the Tethys Sea and the Phylogeny of Eurasian Killifishes (Cyprinodontiformes: Cyprinodontidae). Journal of Evolutionary Biology (J. Evol. Biol.), 16: 1-20, 7 figs., 3 tabs. 

Hrbek, T., C.P. Deus & I.P. Farias. 2004. Rivulus duckensis (Teleostei; Cyprinodontiformes): New Species from the Tarumã Basin of Manaus, Amazonas, Brazil and its Relationship to other Neotropical Rivulidae. Copeia, (3): 569-576, 3 figs., 1 tab.  

Doadrio, I. & O. Dominguez. 2004. Phylogenetic Relationships within the fish Family Goodeidae based on Cytochrome b Sequence data. Molecular Phylogenetics and Evolution (Mol. Phylogenet. Evol.), 31 (2004): 416-430, figs. 

Vermeulen, F.B.M. & T. Hrbek. 2005. Kryptolebias sepia n. sp. (Actinopterygii: Cyprinodontiformes: Rivulidae), a new Killifish from the Tapanahony River drainage in southeast Surinam. Zootaxa, 928: 1-20, 10 figs., 4 tabs.  

Echelle, A.A., E.W. Carson, A.F. Echelle, R.A. Van Den Bussche, T.E. Dowling & A. Meyer. 2005. Historical Biogeography of the New-World Pupfish Genus Cyprinodon (Teleostei: Cyprinodontidae). Copeia, (2): 320-339, 5 figs., 1 tab. 

Hrbek, T., D.C. Taphorn & J.E. Thomerson. 2005. Molecular Phylogeny of Austrofundulus Myers (Cyprinodontiformes: Rivulidae), with Revision of the Genus and the Description of four new species. Zootaxa, 825: 1-39, 16 figs., 9 tabs. 

Legros, O., F. Zentz & J.-F. Agnèse. 2005. Description de deux nouveaux Chromaphyosemion (Teleostei: Aplocheilidae) du Sud de la Plaine littorale camerounaise. Association Killiphile Francophone de Belgique (A.K.F.B.), Killi Contact, 33 (3): 3-30, 13 figs., 1 map, 2 tabs. 

Agnèse, J.-F., F. Zentz, O. Legros & D. Sellos. 2006. Phylogenetic relationships and phylogeography of the Killifish species of the subgenus Chromaphyosemion (Radda, 1971) in West Africa, inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 40 (2), August: 332-346, figs.  

Echelle, A.A., L. Fuselier, R.A. Van den Bussche, C.M.L. Rodriguez & M.L. Smith. 2006. Molecular Systematics of Hispaniolan pupfishes (Cyprinodontidae: Cyprinodon): Implications for the Biogeography of insular Caribbean fishes. Molecular Phylogenetics and Evolution (Mol. Phylogenet. Evol.), 39 (3): 855-864, 3 figs. 

Hrbek, T., Y. Keivany & B.W. Coad. 2006. New Species of Aphanius (Teleostei, Cyprinodontidae) from Isfahan Province of Iran and a Reanalysis of Other Iranian Species. Copeia, (2): 244-255, 4 figs., 5 tabs. 

Sonnenberg, R., T. Blum & B.Y. Misof. 2006. Description of a new Episemion species (Cyprinodontiformes: Nothobranchiidae) from northern Gabon and southeastern Equatorial Guinea. Zootaxa, 1361: 1-20, 5 figs., 1 map, 4 tabs.  

Suijker, W.H. & G.E. Collier. 2006. Rivulus mahdiaensis, a new killifish from central Guyana (Cyprinodontiformes: Rivulidae). Zootaxa, 1246: 1-13, 5 figs., 2 tabs.  

Collier, G.E. 2007. The Genus Aphyosemion: Taxonomic History and Molecular Phylogeny. Journal of the American Killifish Association (J. Amer. Killifish Assoc., JAKA), (2006) 39 (5-6): 147-168, 9 figs.  

Sonnenberg, R. 2007. Description of three new species of the genus Chromaphyosemion Radda, 1971 (Cyprinodontiformes: Nothobranchiidae) from the coastal Plains of Cameroon with a preliminary Review of the Chromaphyosemion splendopleure Complex. Zootaxa, 1591: 1-38. 

Legros, O. & F. Zentz. 2007a. Aphyosemion malumbresi n. sp., un nouveau Chromaphyosemion (Teleostei: Aplocheilidae) originaire de Guinée Equatoriale continentale. Association Killiphile Francophone de Belgique (A.K.F.B.), Killi Contact, (December 2006), 34 (6): 1-28, 4 figs., 1 map, 1 tab. 

Legros, O. & F. Zentz. 2007b. Aphyosemion lividum n. sp. (Cyprinodontiformes: Nothobranchiidae), une nouvelle espèce du secteur d'Edéa au Cameroun. Association Killiphile Francophone de Belgique (A.K.F.B.), Killi Contact, 35 (6) December 2007: 1-34, 4 figs., 2 maps, 2 tabs. 

Hrbek, T. & D.C. Taphorn. 2008. Description of a new annual rivulid Killifish Genus from Venezuela. Zootaxa, 1734: 27-42, figs. 

Agnèse J.-F., R. Brummett, P. Caminade, J. Catalan & E. Kornobis. 2009. Genetic Characterization of the Aphyosemion calliurum species group and Description of a new species from this Assemblage, A. campomaanense (Cyprinodontiformes: Aplocheiloidei: Nothobranchiidae) from southern Cameroon. Zootaxa, 2045: 43-59, 13 figs., 2 tabs. 

Collier, G.E., W.J. Murphy & M. Espinoza. 2009. Phylogeography of the Genus Epiplatys (Aplocheiloidea: Cyprinodontiformes). Molecular Phylogenetics and Evolution, 50 (1): 190-196, figs. 

Sonnenberg, R. & E. Busch. 2009. Description of a new genus and two new species of killifish (Cyprinodontiformes: Nothobranchiidae) from West Africa, with a Discussion of the taxonomic Status of Aphyosemion maeseni Poll, 1941. Zootaxa, 2294: 1-22, 10 figs. 

Collier, G.E. 2010. The Genus Fundulopanchax: Taxonomic History and Molecular Phylogeny. Journal of the American Killifish Association, 43 (1): 3-24, 15 figs. 

Shidlovskiy, K.M., B.R. Watters & R.H. Wildekamp. 2010. Notes on the annual Killifish species Nothobranchius rachovii (Cyprinodontiformes; Nothobranchiidae) with the Description of two new species. Zootaxa, 2724: 37-57. 

Whitehead, A. 2010. The evolutionary Radiation of diverse osmotolerant Physiologies in killifish (Fundulus spp.). Evolution, 64: 2070-2085, 4 figs. 

Costa, W.J.E.M. & P.F. Amorim. 2011. A new annual Killifish species of the Hypsolebias flavicaudatus Complex from the São Francisco River basin, Brazilian Caatinga (Cyprinodontiformes: Rivulidae). Vertebrate Zoology, 61 (1): 99-104, figs. 

Dorn, A., E. Ngoma, K. Janko, K. Reichwald, M. Polacik, M. Platzer, A. Cellerino, M. Reichard. 2011. Phylogeny, genetic variability and colour polymorphism of an emerging animal model: The short-lived annual Nothobranchius fishes from southern Mozambique. Molecular Phylogenetics and Evolution, 61 (2011): 739-749, 2 figs., 6 tabs. 

Sonnenberg, R., T. Woeltjes & J.R. van der Zee. 2011. Description of Fenerbahce devosi (Cyprinodontiformes: Nothobranchiidae), a new species of dwarf Killifish from the eastern Congo Basin in the Democratic Republic of Congo. Zootaxa, 2966: 1-12, figs. 

Zee J.R. van der & R. Sonnenberg. 2011. Aphyosemion musafirii (Cyprinodontiformes: Nothobranchiidae), a new species from the Tshopo Province in the Democratic Republic of Congo, with some Notes on the Aphyosemion of the Congo Basin. Bonn zoological Bulletin, 60 (1): 73-87, figs. 

Costa, W.J.E.M., P.F. Amorim & J.L.O. Mattos. 2012. Species Delimitation in annual killifishes from the Brazilian Caatinga, the Hypsolebias flavicaudatus Complex (Cyprinodontiformes: Rivulidae): Implications for Taxonomy and Conservation. Systematics and Biodiversity, 10 (1): 71-91, figs. 

Garcia G., V. Gutierrez, J. Vergara, P. Calviño, A. Duarte & M. Loureiro. 2012. Patterns of Population Differentiation in annual killifishes from the Parana-Uruguay-La Plata Basin: the Role of Vicariance and Dispersal. Journ. Biogeography, 39: 1707-1719, 6 figs., 2 tabs. 

Matamoros, W.A., Schaefer, J.F., Hernandez, C.L. & P. Chakrabarty. 2012. Profundulus kreiseri, a new species of Profundulidae (Teleostei, Cyprinodontiformes) from northwestern Honduras. Zookeys, 227: 49-62, 4 figs. 

Sonnenberg, R. & E. Busch. 2012. Description of Scriptaphyosemion wieseae (Cyprinodontiformes: Nothobranchiidae), a new species from northern Sierra Leone. Bonn zoological Bulletin, 61 (1): 13-28, 5 figs., 3 tabs.  

Teimori, A., H.R. Esmaeili, Z. Gholami, N. Zarei & B. Reichenbacher. 2012. Aphanius arakensis, a new species of tooth-carp (Actinopterygii, Cyprinodontidae) from the endorheic Namak Lake Basin in Iran. Zookeys, 215: 55-76, figs. 

Valdesalici, S., R. Bills, A. Dorn, K. Reichwald & A. Cellerino. 2012. Nothobranchius niassa (Cyprinodontiformes: Nothobranchiidae), a new species of annual Killifish from northern Mozambique. Ichthyological Exploration of Freshwaters, 23 (1): 19-28, 5 figs., 3 tabs. 

Agnèse, J.-F., O. Legros, B. Cazaux & G. Estivals. 2013. Aphyosemion pamaense, a new killifish species (Cyprinodontiformes: Nothobranchiidae) from Cameroon. Zootaxa, 3670 (4): 516-530, 10 figs., 2 tabs.  

Esquer, G.Y., B. Hugueny, K. Koerner, C. Ibanez, C. Bonillo, P. Pruvost, R. Causse, C. Cruaud & P. Gaubert. 2013. Non-invasive ancient DNA protocol for fluid-preserved specimens and phylogenetic systematics of the genus Orestias (Teleostei: Cyprinodontidae). Zootaxa, 3640 (3): 373-394, 5 figs., 5 tabs.  

Ngoma, E., S. Valdesalici, K. Reichwald & A. Cellerino. 2013. Genetic and morphological Studies of Nothobranchius (Cyprinodontiformes) from Malawi with Description of Nothobranchius wattersi sp. nov. Journal of Fish Biology, 82 (1): 165-188, 5 figs., 6 tabs. 

Vila I., P. Morales, S. Scott, E. Poulin, D. Véliz, C. Harrod & M.A. Mendez. 2013. Phylogenetic and phylogeographic Analysis of the genus Orestias (Teleostei: Cyprinodontidae) in the southern Chilean Altiplano: the Relevance of ancient and recent Divergence Processes in Speciation. Journal of Fish Biology, 82: 927-943, 3 figs. 

Wildekamp, R.H., B.R. Watters & K.M. Shidlovskiy. 2014. Review of the Nothobranchius neumanni species group with descriptions of three new species from Tanzania (Cyprinodontiformes: Nothobranchiidae). Journal of the American Killifish Association, 47 (1) : 2-30, 23 figs. 

Pfleiderer, S.J., M.F. Geiger & F. Herder. 2014. Aphanius marassantensis, a new toothcarp from the Kizilirmak drainage in northern Anatolia (Cyprinodontiformes: Cyprinodontidae). Zootaxa, 3887 (5): 569-582, 8 figs. 

Dorn, A., Z. Musilova, M. Platzer, K. Reichwald & A. Cellerino. 2014. The strange Case of East African annual fish: Aridification correlates with Diversification for a Savannah aquatic Group? BMC Evolutionary Biology, 14: 210-235, figs., tabs., suppl. 

Gholami, Z., H.R. Esmaeili, D. Erpenbeck & B. Reichenbacher. 2014. Phylogenetic analysis of Aphanius from the endorheic Kor River Basin in the Zagros Mountains, South-western Iran (Teleostei: Cyprinodontiformes: Cyprinodontidae). Journal of Zoological Systematics and Evolutionary Research, 52 (2) (May): 130-141. 

Freyhof, J., H. Kärst & M. Geiger. 2014. Valencia robertae, a new killifish from southern Greece (Cyprinodontiformes: Valenciidae). Ichthyological Exploration of Freshwaters (Ichthyol. Explor. Freshwaters), 24 (4) (2013): 289-298, 11 figs. 

Costa, W.J.E.M., P.F. Amorim & G.N. Aranha. 2014. Species Limits and DNA barcodes in Nematolebias, a genus of seasonal killifishes threatened with Extinction from the Atlantic Forest of south-eastern Brazil, with description of a new species (Teleostei: Rivulidae). Ichthyological Exploration of Freshwaters, 24 (3) (2013): 225-236, 3 figs., 2 tabs. 

Garcia, G., V. Gutiérrez, N. Rios, B. Turner, F. Santinaque, B. Lopez-Carro & G. Folle. 2014. Burst speciation processes and genomic expansion in the neotropical annual killifish genus Austrolebias (Cyprinodontiformes, Rivulidae). Genetica, 142 (1): 87-98, 4 figs. 

Costa, W.J.E.M., P.F. Amorim & P.H.N. Bragança. 2014. Species Limits and phylogenetic Relationships of red-finned cryptic species of the seasonal killifish genus Hypsolebias from the Brazilian semi-arid Caatinga (Teleostei: Cyprinodontiformes: Rivulidae). Journal of Zoological Systematics and Evolutionary Research, 52 (1) (February): 52-58, figs. 

Pollux, B.J.A., R.W. Meredith, M.S. Springer, T. Garland & D.N. Reznick. 2014. The Evolution of the Placenta drives a Shift in sexual Selection in livebearing fish. Nature, 513: 233-236, figs., Suppl. 

Pohl, M., F.C. Milvertz, A. Meyer & M. Vences. 2015. Multigene Phylogeny of cyprinodontiform fishes suggests continental Radiations and a rogue Taxon Position of Pantanodon. Vertebrate Zoology, 65 (1): 37-44, 6 figs., 2 tabs. 

Takahashi, T. & E. Moreno. 2015. A RAD-based Phylogenetics for Orestias fishes from Lake Titicaca. Molecular Phylogenetics and Evolution, 93 (December): 307-317. 

Morcillo, F., C.P. Ornelas-Garcia, L. Alcaraz, W.A. Matamoros & I. Doadrio. 2016. Phylogenetic Relationships and Evolutionary History of the Mesoamerican endemic Freshwater fish family Profundulidae (Cyprinodontiformes: Actinopterygii). Molecular Phylogenetics and Evolution, 94, Part A (January 2016): 242-251, 5 figs.  

Costa, W.J.E.M., Amorim, P.F. & J.L.O. Mattos. 2016. Molecular Phylogeny and Evolution of internal Fertilization in South American seasonal cynopoeciline killifishes. Molecular Phylogenetics and Evolution, 95 (February): 94-99, figs. 

Ponzetto, J.M., R. Britzke, D.T.B. Nielsen, P.P. Parise-Maltempi & A.L. Alves. 2016. Phylogenetic Relationships of Simpsonichthys subgenera (Cyprinodontiformes, Rivulidae), including a Proposal for a new genus. Zoologica Scripta, DOI: 10.1111/zsc.12159 

Costa, W.J.E.M, P.F. Amorim & J.L. Mattos. 2016. A new species of inseminating seasonal killifish of the Cynopoecilus melanotaenia complex from southern Brazil (Cyprinodontiformes: Rivulidae). Biodiversity Data Journal, 4: e6888, 4 figs., 2 tabs.  

Britzke, R., D.T.B. Nielsen & C. Oliveira. 2016. Description of two new species of annual fishes of the Hypsolebias antenori species group (Cyprinodontiformes: Rivulidae), from Northeast Brazil. Zootaxa, 4114 (2): 123-138, 7 figs., 2 tabs.  

Costa, W.J.E.M. 2016a. Inferring Evolution of Habitat Usage and Body Size in endangered, seasonal Cynopoeciline Killifishes from the South American Atlantic Forest through an integrative Approach (Cyprinodontiformes: Rivulidae). PLoS ONE, 1-16, 2 figs, 2 tabs., 2 appx. 

Jamangapé, O.J.A., E. Velazquez-Velazques, E. Marínez-Ramirez, M.J. Anzueto-Calvo, E.L. Gomez, S.E. Dominguez-Cisneros, C.D. McMahan & W.A. Matamoros. 2016. Validity and Redescription of Profundulus balsanus Ahl, 1935 (Cyprinodontiformes: Profundulidae). Zootaxa, 4073 (1): 55-65. 



Note : a special note of thanks to Alexander Dorn for valuable comments and improvements on this table.